Lousiana State University AgCenter Entomology Department - Research

NSF Grants:

Schowalter, T.D. and H.K. Luh. Canopy arthropod responses to storm disturbances at U.S. and Taiwanese LTER sites. NSF International Programs, INT-0107485, 2001-04, $74,263.

Lowman, M.D., M.D. Hunter and T.D. Schowalter. Canopy herbivory and soil processes in a temperate and tropical forest. NSF Ecosystem Studies, DEB-9815133, 1999-04, $300,000 (Schowalter $71,559).

Schowalter, T.D. and J.J. Morrell. Heterotroph and moisture effects on log decomposition processes. NSF Ecosystem Studies, DEB-9408169, 1994-99, $500,903. REU Supplements, 1995-99, $45,000.

Swanson, F.J. et al. Long-term ecological research at the H.J. Andrews Experimental Forest. NSF Long Term Ecological Research, BSR-9011663, 1991-96, Schowalter, $60,000.

Waide, R.B. et al. Long-term ecological research on the Luquillo Experimental Forest II. NSF Long Term Ecological Research, 1994-2000, Schowalter, $60,000.

Schowalter, T.D., J.D. Lattin, S.E. Carpenter, R.G. Kelsey, E.R. Ingham, M.E. Harmon, R.P. Griffiths, and B.A. Caldwell. Decomposition processes as influenced by heterotroph community structure. NSF Ecosystem Studies, BSR-8717434, 1988-89, $50,000.

Myrold, D.D., P.A. Wheeler, H.J. Evans, R. Molina, D.A. Perry, K. Cromack, Jr., and T.D. Schowalter. Acquisition of an automated ration mass spectrometer for ¹⁵N and ¹³C analysis. NSF Biological Instrumentation, DIR 8813197, 1988, $62,323.

Schowalter, T.D., J.D. Lattin, J.M. Trappe, S.E. Carpenter, and M.E. Harmon. Heterotroph effects on the log decomposition process in a coniferous forest ecosystem. NSF Ecosystem Studies, BSR-8516590, 1986-87, $425,000.

Schowalter, T.D. Phytophagous arthropod effects on nutrient loss and decomposition of Douglas-fir foliage at the H.J. Andrews Experimental Forest, Oregon. NSF Ecosystem Studies, BSR-8306490, 1983-86, $180,000.

Oak Ridge Assoc. Univ. (DOE) Contracts:

Schowalter, T.D. Canopy invertebrate responses to Hurricane Hugo in a tropical rainforest, Puerto Rico. Oak Ridge Associated Universities Travel Contract S-3441, 1991-98, $9,000.

EPA Contracts:

Schowalter, T.D. Development of a protocol for testing effects of microbial pest control agents on non-target litter microarthropods. U.S.E.P.A., 1988-89, $1,000.

USDA Forest Service Cooperative Agreements:

Schowalter, T.D. Abundance, diversity, and chemical ecology of Alien Invasive Ambrosia Beetles in Central Louisiana. Southern Forest Res. Stn., SRS 04-CA-11330129-059, 2004-06, $36,092.

Schowalter, T.D. Litter invertebrate responses to green tree retention treatments in the Demonstration of Ecosystem Management Options study. Pacific Northwest Res. Stn., PNW 02-CA-11261993-104, 2002-2005, $147,668 (relinquished on departure from OSU).

Schowalter, T.D. Litter invertebrate response to forest thinning and woody debris removal. Pacific Northwest Res. Stn., 98-9048-2CA PNW, 1999-2001, $40,000.

Schowalter, T.D. Canopy invertebrates in mixed-conifer old-growth at Teakettle Experimental Forest. Pacific Southwest Res. Stn., PSW-98-010-RJVA, 1998-03, $106,100.

Schowalter, T.D. Effects of silvicultural treatments on composition and function of coarse woody debris decomposers. Pacific Southwest Res. Stn. PSW 98-0008CA, 1998-2001, $82,000.

Schowalter, T.D. Invertebrate responses to alternative forest thinning practices. Pacific Northwest Res. Stn. D.E.M.O. Project, PNW 96-5012-55-CA, 1996-2001, $234,500.

Schowalter, T.D. Effect of resource concentration among site productivity treatments on insect abundance and effects on plant growth and survival. Pacific Northwest Res. Stn., requested $40,000, granted $8,000 for 1992-93, declined due to departure to NSF.

Schowalter, T.D. Effects of thinning on southern pine beetle risk to old-growth stands. Southern Forest Exp. Stn., 19-92-031, 1992-94, $7,000, declined due to departure to NSF.

Schowalter, T.D. Evaluation of burning to reduce overwintering populations of gall midge pupae and seed chalcid larvae in Douglas-fir seed orchards. Pacific Northwest Res. Stn., 1991-95, $49,799.

Schowalter, T.D. Resource concentration thresholds for southern pine beetle infestation development. Southern Forest Exp. Stn., 19-89-037, 1989-91, $84,475.

Schowalter, T.D. Seasonal history and infestation characteristics of the balsam woolly adelgid in selected areas in northern Idaho. PNW 90-501, 1990-92, $13,948.

Schowalter, T.D. Management of cone and seed insects of western white pine and Douglas-fir from seed orchards in western Oregon. PSW-89-0009CA, 1989-91, $16,000.

Kelsey, R.G., T.D. Schowalter, J.C. Miller, and G.D. Daterman. Isolation and identification of gypsy moth feeding deterrents from plant sources. PNW-87-409, 1987-89, $72,186.

Schowalter, T.D. Determination of the annual variation in the distribution of damage caused by cone and seed insects in Douglas-fir seed orchards in California, Oregon, and Washington, and assessment of the role of geography, genetics, and land-use. PSW-84-0023CA, 1984-86, $12,500.

Schowalter, T.D. Development of a model for predicting impact of Douglas-fir cone midge on seed production in Douglas-fir seed orchards. PSW-83-0010CA, 1983-86, $13,000.

Schowalter, T.D. Field survey to identify the agent and establish the distribution and abundance of Douglas-fir tip die-back in Douglas-fir nurseries. PSW-84-0012CA, 1984-85, $8,358.

Schowalter, T.D. Determination of the distribution and severity of damage caused by cone and seed insects in Douglas-fir seed orchards in California, Oregon, and Washington. PSW-83-0023CA, 1983-84, $6,000.

Professor, Department of Entomology, Louisiana State University AgCenter, 2015-present

Professor and Head, Department of Entomology, Louisiana State University AgCenter, 2003-2015

Interim Head, Department of Entomology, Oregon State University, 2001-03

Professor 1993-2003, Associate Professor 1987-93, Assistant Professor 1981-87, Department of Entomology, Oregon State University, 1993-present.

Program Director for Ecological Studies, National Science Foundation, 1992-93.

Visiting Associate Professor, Department of Biology, New Mexico State University, 1990-91.

Postdoctoral Fellow, Department of Entomology, Texas A & M University, 1979-81.

Other Professional Experience:

Associate Editor, Population Ecology, 2007-present

Associate Editor, Frontiers in Ecology and the Environment, 2005-present

Chair, Council of Entomology Department Administrators, 2009

Co-organizer of Council of Entomology Department Administrators (CEDA)-sponsored symposium on Metamorphosis of Entomology into a Cornerstone Discipline in Social and Environmental Issues for Entomological Society of America Meeting, Reno, NV, November 2008.

Chair-elect, Council of Entomology Department Administrators, 2008

Invited participant, Ecological Society of America/U.S. Department of Defense workshop on pine forest decline in the southeastern U.S., Atlanta, GA., June 2007.

Invited participant, National Commission on Science and Sustainable Forestry symposium on Forest Disturbance, Management and Biodiversity, Denver, CO, April 2006.

Elected Secretary, AC-12, Southern Entomology Administrators Group, 2004-05; rotated to Chair, 2006-07.

Elected Vice President for Public Affairs and Governing Board member, Ecological Society of America, 1999-2002.

Member, Long Term Ecological Research Program delegation to Taiwan to develop collaborative arrangements among U.S. and Taiwanese LTER sites, March 2000.

Invited participant, National Science Foundation workshop to develop guidelines for Biological Observatories Network (BON) Program, National Center for Ecological Analysis and Synthesis, Santa Barbara, CA, August, 1999.

National Academy of Sciences, National Research Council Committee on Environmental Issues in Pacific Northwest Forest Management, 1993-99.

Co-Organizer, “Canopy-forest floor interactions” session, International Canopy Symposium, Sarasota, FL, November, 1998.

Organizer, workshop on “Insects as agents of ecosystem change”, Western Forest Insect Work Conference, Jackson, WY, March, 1998.

Co-organizer, discussion session on “Management of pest resistance in transgenic crops expressing the Bt toxin, held at the Ecological Society of America Annual Meeting, Albuquerque, NM, August, 1997.

Panelist, USDA NRICGP Forest/Range/Crop/Aquatic Ecosystems Program, 1996.

Governor's Working Group on East Side Forest Health, 1995-present.

Member, Long Term Ecological Research Program delegation to Taiwan International LTER Conference to develop collaborative arrangements among east Asia and Pacific nations and U.S. LTER sites, 1995.

National Scientific Advisory Committee, Wind River Canopy Crane, 1994-present.

Presider, Apprenticeships in Science and Engineering Symposium, Salem, OR, August 1994, Corvallis, OR, August 1995.

Public Affairs Committee, Ecological Society of America, 1994-97.

Member, Long Term Ecological Research Program delegation to Hungary to develop collaborative arrangements with U.S. LTER sites, 1994-95.

Member, panel to review the Savannah River Site Biodiversity Research Program, 1994.

NSF Representative, UCAR Office of Interdisciplinary Studies Global Change Institute, Snowmass, CO, July 1993.

NSF Representative, UNCED Convention on Desertification Policy and Technical Task Forces, 1992-93.

NSF Representative (alternate), Presidential Review Directive, Interagency Desertification Task Force, 1993

NSF Representative, Terrestrial Ecosystem Initiative Panel, Process Working Group, Subcommittee on Global Change Research (SGCR), Committee on Earth and Environmental Sciences (CEES), Federal Coordinating Committee for Science, Engineering and Technology (FCCSET), 1993

Refereed Journal Articles:

Schowalter, T.D. 2012. Insect herbivore effects on forest ecosystem services. Journal of Sustainable Forestry (in press).

Schowalter, T.D. 2012. Ecology and management of bark beetles (Coleoptera: Curculionidae: Scolytinae) in southern pine forests. Journal of Integrated Pest Management 3: A1-7.

Schowalter, T.D., S.J. Fonte, J. Geaghan and J. Want. 2011. Effects of manipulated herbivore inputs on nutrient flux and decomposition in a tropical rainforest in Puerto Rico. Oecologia 167: 1141-1149.

Schowalter, T.D. 2008. Insect Herbivore Responses to Management Practices in Conifer Forests in North America. Journal of Sustainable Forestry 26: 204-222.

Fonte, S.J. and T.D. Schowalter. 2005. The influence of a neotropical herbivore (Lamponius portoricensis) on nutrient cycling and soil processes. Oecologia 146: 423-431.

Schowalter, T.D. and Y.L. Zhang. 2005. Canopy arthropod assemblages in four overstory and three understory plant species in mixed-conifer old-growth forest in California. Forest Science 51: 233-242.

Schowalter, T.D., Y.L. Zhang and R.A. Progar. 2005. Canopy arthropod response to density and distribution of green trees retained after partial harvest. Ecological Applications 15: 1594-1603.

Fonte, S.J. and T.D. Schowalter. 2004. Decomposition of greenfall vs. senescent foliage in relation to herbivory and disturbance in a forest ecosystem. Biotropica 36: 374-382.

Schowalter, T.D., Y.L. Zhang and J.J. Rykken. 2003. Litter invertebrate responses to variable density thinning in western Washington forest. Ecological Applications 13: 1204-1211.

Progar, R.A. and T.D. Schowalter. 2002. Canopy arthropod assemblages along a precipitation and latitudinal gradient among Douglas-fir Pseudotsuga menziesii forests in the Pacific Northwest of the United States. Ecography 25: 129-138.

Schowalter, T.D. and J.J. Morrell. 2002. Nutritional quality of Douglas-fir wood: effect of vertical and horizontal position on nutrient levels. Wood and Fiber Science 34: 158-164.

Schowalter, T.D. and J. Withgott. 2001. Rethinking insects: what would an ecosystem approach look like? Conservation Biology in Practice 2: 10-16.

Smith, J.P. and T.D. Schowalter. 2001. Aphid-induced reduction in shoot and root growth of Douglas-fir seedlings. Ecological Entomology 26: 411-416.

Progar, R., T.D. Schowalter, J.J. Morrell and C.M. Freitag. 2000. Respiration from coarse woody debris as affected by moisture and saprotroph functional diversity in western Oregon. Oecologia 124: 426-431.

Schowalter, T.D., D.C. Lightfoot and W.G. Whitford. 1999. Diversity of arthropod responses to host plant water stress in southern New Mexico. American Midland Naturalist 142: 281-290.

Schowalter, T.D. and L.M. Ganio. 1999. Invertebrate communities in a tropical rain forest canopy in Puerto Rico following Hurricane Hugo. Ecological Entomology 24: 191-201.

Progar, R.A., T.D. Schowalter and T. Work. 1999. Arboreal invertebrate responses to varying levels and patterns of green-tree retention in northwestern forests. Northwest Science 73 (spec. issue): 77-86

Schowalter, T.D. 1999. Throughfall volume and chemistry as affected by precipitation volume, sapling size, and canopy herbivory in a regenerating Douglas-fir ecosystem in western Oregon. Great Basin Naturalist 59: 79-84.

Schowalter, T.D. and L.M. Ganio. 1998. Vertical and seasonal variation in canopy arthropod abundances in an old-growth conifer forest in southwestern Washington. Bulletin of Entomological Research 88: 633-640.

Mankowski, M.E., T.D. Schowalter, J.J. Morrell and B.Lyons. 1998. Feeding habits and gut fauna of Zootermopsis angusticollis (Isoptera: Termopsidae) in response to wood species and fungal associates. Environmental Entomology 27: 1315-1322.

Goyer, R.A., M.R. Wagner and T.D. Schowalter. 1998. Current and proposed technologies for bark beetle management. Journal of Forestry 97(12): 29-33.

Schowalter, T.D., Y.L. Zhang and T.E. Sabin. 1998. Decomposition and nutrient dynamics of oak (Quercus spp.) logs after five years of decomposition. Ecography. 21: 3-10.

Zhang, Y. and T.D. Schowalter. 1997. Cone and seed insect management in seed orchards: an example from Douglas-fir. Journal of Forestry 95(3): 28-32.

Schowalter, T.D. 1996. Arthropod associates and herbivory on tarbush in southern New Mexico. Southwestern Naturalist 41: 140-44.

Schowalter, T.D. 1995. Canopy arthropod community responses to forest age and alternative harvest practices in western Oregon. Forest Ecology and Management 78: 115-25.

Schowalter, T.D. 1995. Canopy invertebrate community response to disturbance and consequences of herbivory in temperate and tropical forests. Selbyana 16: 41-8.

Schowalter, T.D. 1994. Cone and seed insect phenology in a Douglas-fir seed orchard during three years in western Oregon. Journal of Economic Entomology 87:758-65.

Schowalter, T.D. 1994. Invertebrate community structure and herbivory in a tropical rainforest canopy in Puerto Rico following Hurricane Hugo. Biotropica 26:312-19

Schowalter, T.D. and P. Turchin. 1993. Southern pine beetle infestation development: interaction between pine and hardwood basal areas. Forest Science 39:201-10.

Franklin, J.F., F.J. Swanson, M.E. Harmon, D.A. Perry, T.A. Spies, V.H. Dale, A. McKee, W.K. Ferrell, J.E. Means, S.V. Gregory, J.D. Lattin, T.D. Schowalter and D. Larsen. 1992. Effects of global climatic change on forests in northwestern North America. Northwest Environmental Journal 7:233-54.

Schowalter, T.D. 1992. Early decomposition and nutrient dynamics of oak (Quercus) logs at four sites across a North American gradient. Canadian Journal of Forest Research 22:161-66.

Schowalter, T.D. and T.E. Sabin. 1991. Litter microarthropod response to canopy herbivory, season and decomposition in litterbags in a regenerating conifer ecosystem in western Oregon. Soil Biology and Fertility 11:93-96.

Sexton, J.M. and T.D. Schowalter. 1991. Physical barriers to reduce Lepesoma lecontei (Coleoptera: Curculionidae) damage to conelets in a Douglas-fir seed orchard in western Oregon. Journal of Economic Entomology 84:212-14.

Connelly, A.E. and T.D. Schowalter. 1991. Seed losses to feeding by Leptoglossus occidentalis (Heteroptera: Coreidae) during two periods of second year cone development in western white pine. Journal of Economic Entomology 84:215-17.

Schowalter, T.D., T.E. Sabin, S.G. Stafford, and J.M. Sexton. 1991. Phytophage effects on primary production, nutrient turnover, and litter decomposition of young Douglas-fir in western Oregon. Forest Ecology and Management 42:229-43.

Schowalter, T.D. and J.M. Sexton. 1990. Effect of Leptoglossus occidentalis (Heteroptera: Coreidae) on seed development of Douglas-fir at different times during the growing season in western Oregon. Journal of Economic Entomology 83:1485-86.

Zhong, H. and T.D. Schowalter. 1989. Conifer bole utilization by wood-boring beetles in western Oregon. Canadian Journal of Forest Research 19:943-47.

Schowalter, T.D. 1989. Canopy arthropod community structure and herbivory in old-growth and regenerating forests in western Oregon. Canadian Journal of Forest Research 19:318-22.

Schowalter, T.D. and M.I. Haverty. 1989. Influence of host genotype on Douglas-fir seed losses to Contarinia oregonensis (Diptera: Cecidomyiidae) and Megastigmus spermotrophus (Hymenoptera: Torymidae) in western Oregon. Environmental Entomology 18:94-97.

Schowalter, T.D., S.G. Stafford, and R.L. Slagle. 1988. Arboreal arthropod community structure in an early successional coniferous forest ecosystem in western Oregon. Great Basin Naturalist 48:327-33.

Dombrosky, S.A. and T.D. Schowalter. 1988. Inventory monitoring for estimation of impact of insects on seed production in a Douglas-fir seed orchard in western Oregon. Journal of Economic Entomology 81:281-85.

Schowalter, T.D. 1988. Forest pest management: a synopsis. Northwest Environmental Journal 4:313-18.

Witcosky, J.J., T.D. Schowalter, and E.M. Hansen. 1987. Host-derived attractants for the beetles, Hylastes nigrinus (Coleoptera: Scolytidae) and Steremnius carinatus (Coleoptera: Curculionidae). Environmental Entomology 16:1310-13.

Schowalter, T.D. and J.D. Stein. 1987. Influence of Douglas-fir seedling provenance and proximity to insect population sources on susceptibility to Lygus hesperus (Heteroptera: Miridae) in a forest nursery in western Oregon. Environmental Entomology 16:984-86.

Schowalter, T.D. 1987. Abundance and distribution of Lygus hesperus (Heteroptera: Miridae) in two conifer nurseries in western Oregon. Environmental Entomology 16:687-90.

Witcosky, J.J., T.D. Schowalter, and E.M. Hansen. 1986. Hylastes nigrinus (Coleoptera: Scolytidae) and Pissodes fasciatus and Steremnius carinatus (Coleoptera: Curculionidae) as vectors of black-stain root disease of Douglas-fir. Environmental Entomology 15:1090-95.

Witcosky, J.J., T.D. Schowalter, and E.M. Hansen. 1986. The influence of time of precommercial thinning on the colonization of Douglas-fir by three species of root-colonizing insects. Canadian Journal of Forest Research 16:745-49.

Schowalter, T.D. 1986. Overwintering aggregation of Boisea rubrolineatus (Heteroptera: Rhopalidae) in western Oregon. Environmental Entomology 15:1055-56.

Schowalter, T.D. 1986. Ecological strategies of forest insects: the need for a community-level approach to reforestation. New Forests 1:57-66.

Schowalter, T.D., D.L. Overhulser, A. Kanaskie, J.D. Stein, and J. Sexton. 1986. Lygus hesperus as an agent of apical bud abortion in Douglas-fir nurseries in western Oregon. New Forests 1:5-15.

Schowalter, T.D., M.I. Haverty, and T.W. Koerber. 1985. Cone and seed insects in Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) seed orchards in the western United States: distribution and relative impact. Canadian Entomologist 117:1223-30.

Schowalter, T.D. 1984. Comparison of arthropods emerging in the spring from Douglas-fir litter between a mature stand and a seed orchard in western Oregon. Environmental Entomology 13:1253-55.

Schowalter, T.D. 1984. Dispersal of cone and seed insects to an isolated Douglas-fir tree in western Oregon. Canadian Entomologist 116:1437-38.

Schowalter, T.D. and D.A. Crossley, Jr. 1983. Forest canopy arthropods as sodium, potassium, magnesium and calcium sinks in forests. Forest Ecology and Management 7:143-48.

Schowalter, T.D., R.N. Coulson, R.H. Turnbow, and W.S. Fargo. 1982. Accuracy and precision of procedures for surveying populations of the southern pine beetle, Dendroctonus frontalis (Coleoptera: Scolytidae). Journal of Economic Entomology 75:1009-16.

Fargo, W.S., T.L. Wagner, R.N. Coulson, J.D. Cover, T. McAudle, and T. D. Schowalter. 1982. Probability functions for components of the Dendroctonus frontalis-host tree population system and their potential use with population models. Researches in Population Ecology 24:123-31.

Schowalter, T.D. and D.A. Crossley, Jr. 1982. Bioelimination of 51Cr and 85Sr by cockroaches, Gromphadorhina portentosa (Orthoptera: Blaberidae), as affected by mites, Gromphadorholaelaps schaeferi (Parasitiformes: Laelapidae). Annals of the Entomological Society of America 75:158-60.

Schowalter, T.D., D.N. Pope, R.N. Coulson, and W.S. Fargo. 1981. Patterns of southern pine beetle (Dendroctonus frontalis Zimm.) infestation enlargement. Forest Science 27:837-49.

Schowalter, T.D., R.N. Coulson, and D.A. Crossley, Jr. 1981. Role of southern pine beetle and fire in maintenance of structure and function of the southeastern coniferous forest. Environmental Entomology 10:821-25.

Schowalter, T.D., J.W. Webb, and D.A. Crossley, Jr. 1981. Community structure and nutrient content of canopy arthropods in clearcut and uncut forest ecosystems. Ecology 62:1010-19.

Schowalter, T.D. 1981. Insect herbivore relationships to the state of the host plant: biotic regulation of ecosystem nutrient cycling through ecological succession. Oikos 37:126-30.

Schowalter, T.D. and W.G. Whitford. 1979. Territorial behavior of Bootettix argentatus Bruner (Orthoptera: Acrididae). American Midland Naturalist 102:182-84.

Schowalter, T.D. and M.J. Klowden. 1979. Blood meal size of the stable fly, Stomoxys calcitrans, measured by the HiCN method. Mosquito News 29:110-12.

Schowalter, T.D., W.G. Whitford, and R.B. Turner. 1977. Bioenergetics of the range caterpillar, Hemileuca oliviae (Ckll.). Oecologia 28:153-61.

Schowalter, T.D., W.G. Whitford, and R.B. Turner. 1976. Growth characteristics of the range caterpillar, Hemileuca oliviae Ckll.) on an artificial diet. Southwestern Entomologist 1:164-67.

Review Articles:

Lowman, M.D. and T.D. Schowalter. 2012. Tansley Review: Plant science in forest canopies - the first forty years of advances and challenges (1980-2010). New Phytologist 194: 12-27.

Schowalter, T.D. 2012. Insect responses to major landscape-level disturbance. Annual Review of Entomology 57: 1-20.

Schowalter, T.D., W.W. Hargrove, and D.A. Crossley, Jr. 1986. Herbivory in forested ecosystems. Annual Review of Entomology 31:177-96.

Books and Book Chapters:

Schowalter, T.D. Insects and the Sustainability of Ecosystem Services. Taylor and Francis (in prep.)

Lowman, M.D., T.D. Schowalter and J.F. Franklin. 2012. Methods in Forest Canopy Research. University of California Press (in press).

Schowalter, T.D. 2012. Insect outbreak effects on ecosystem services. In P. Barbosa, D.K. Letourneau and A.A. Agrawal, eds., Insect Outbreaks Revisited. Wiley/Blackwell (in press).

Brokaw, N.V., J.K. Zimmerman, M.R. Willig...T.D. Schowalter, et al. 2012. Response to disturbance. pp. 201-271 In N.V. Brokaw, T.A. Crowl, A.E. Lugo et al., eds., A Caribbean Tapestry: the Multidimensional Nature of Disturbance and Response. Oxford University Press, New York.

Schowalter, T.D. 2011. Insect Ecology: an Ecosystem Approach, 3rd Ed. Elsevier/Academic, San Diego, CA.

Schowalter, T.D. 2007. Ecosystems: concepts, analyses and practical implications, Pp. 411-430, In: M. Kogan and P. Jepson, eds., Perspectives in Ecological Theory and Integrated Pest Management. Cambridge University Press, Cambridge, U.K.

Schowalter, T.D. 2006. Insect Ecology: an Ecosystem Approach, 2^nd Ed. Elsevier/Academic, San Diego, CA.

Fonte, S.J. and T.D. Schowalter. 2004. Decomposition in forest canopies. pp. 413-422, In: M.D. Lowman and B. Rinker, eds., Forest Canopies. Elsevier, Amsterdam.

Schowalter, T.D. and L.M. Ganio. 2003. Diel, seasonal and disturbance-induced variation in invertebrate assemblages. pp. 315-328, In: Y. Basset, V. Navotny, S. Miller and R. Kitching, eds., Arthropods of Tropical Forests. Cambridge University Press, Cambridge, UK.

Christensen, N.L., Jr., S.V. Gregory, P.R. Hagenstein, T.A. Heberlein, J.C. Hendee, J.T. Olson, J.M. Peek, D.A. Perry, T.D. Schowalter, K. Sullivan, G.D. Tilman and K.A. Vogt. 2000. Environmental Issues in Pacific Northwest Forest Management. National Academy Press, Washington, D.C.

Schowalter, T.D. 2000. Insect Ecology: an Ecosystem Approach. Academic Press, San Diego, 483 pp.

Schowalter, T.D. and M.D. Lowman. 1999. Forest herbivory. pp. 269-285, In: L.R. Walker (Ed.) Ecosystems of the World: Ecosystems of Disturbed Ground. Elsevier, Amsterdam.

Schowalter, T.D. E.M. Hansen, R.L. Molina and Y.L. Zhang. 1996. Integrating the ecological roles of phytophagous insects, plant pathogens, and mycorrhizae in managed forests. pp. 171-189, In: K.A. Kohm and J.F. Franklin (Eds.) Creating a Forestry for the 21st Century: The Science of Ecosystem Management. Island Press, Washington, D.C.

Schowalter, T.D. and G.M. Filip, eds. 1993. Beetle-Pathogen Interactions in Conifer Forests. Academic Press, London. 252 pp.

Schowalter, T.D. and G.M. Filip. 1993. Bark beetle-pathogen-conifer interactions: an overview. pp. 3-19, In: T.D. Schowalter and G.M. Filip (Eds.), Beetle-Pathogen Interactions in Conifer Forests. Academic Press, London.

Filip, G.M. and T.D. Schowalter. 1993. Synopsis. pp. 231-238, In: T.D. Schowalter and G.M. Filip (Eds.), Beetle-Pathogen Interactions in Conifer Forests. Academic Press, London.

Helgerson, O.T., M. Newton, D. deCalesta, T.D. Schowalter, and E.M. Hansen. 1992. Protecting young regeneration. pp. 384-420, In: S.D. Hobbs, S.D. Tesch, P.W. Owston, R.E. Stewart, J.C. Tappeiner II, and G.E. Wells (Eds.), Reforestation Practices in Southwestern Oregon and Northern California. Forest Research Lab, Oregon State University, Corvallis.

Schowalter, T.D. 1990. Lygus bugs. pp. 50-52, In: P.B. Hamm, S.J. Campbell, and E.M. Hansen (Eds.), Growing Healthy Seedlings: Identification and Management of Pests in Northwest Forest Nurseries. Spec. Publ. 19, Forest Research Lab, Oregon State University, Corvallis.

Schowalter, T.D. 1985. Adaptations of insects to disturbance. pp. 235-52, In: Pickett, S.T.A. and P.S. White (Eds.) The Ecology of Natural Disturbance and Patch Dynamics. Academic Press, New York.

Symposia Proceedings:

Schowalter, T.D. 2000. Insects as regulators of ecosystem development. pp. 99-114, In D.C. Coleman and P.F. Hendrix (Eds.) Invertebrates as Webmasters in Ecosystems. CABI Publishing, Wallingford, U.K.

Schowalter, T.D. 1996. Stand and landscape diversity as a mechanism of forest resistance to insects. pp. 21-27, In: W.J. Mattson, P. Niemela and M. Rousi (Eds.) Dynamics of Forest Herbivory: Quest for Pattern and Principle. USDA Forest Serv. Gen. Tech. Rpt. NC-183. USDA Forest Serv. North Central Forest Exp. Stn., St. Paul, MN.

Schowalter, T.D. 1994. Primary productivity as affected by insect herbivores. pp. 219-227, In: P. Singh, P.S. Pathak, V. Shankar and A.K. Srivastava (Eds.), Rangeland Resources - Management and Utilization, Proc. Third International Rangeland Congress, 7-11 November 1988, New Delhi, India, Range Management Society of India, Jhansi, India.

Schowalter, T.D. 1994. An ecosystem-centered view of insect and disease effects on forest health. pp. 189-195, In: W.W. Covington and L.F. DeBano, (Eds.), Sustainable ecological systems: implementing an ecological approach to land management. USDA Forest Serv. Gen. Tech. Rep. RM-247. USDA Forest Serv., Rocky Mountain Forest & Range Exp. Stn., Fort Collins, CO.

Schowalter, T.D. 1992. Insect contributions to nutrient cycling and biodiversity in regenerating forests. p. 116, In: D.C. Allen and L.P. Abrahamson (Eds.), Proc. North American Forest Insect Work. Conf., USDA Forest Serv. Gen. Tech. Rpt. PNW-GTR-294. USDA Forest Serv. Pacific Northwest Res. Stn., Portland OR.

Franklin, J.F., F.J. Swanson, M.E. Harmon, D.A. Perry, T.A. Spies, V.H. Dale, A. McKee, W.K. Ferrell, J.E. Means, S.V. Gregory, J.D. Lattin, T.D. Schowalter and D. Larsen. 1992. Effects of global climatic change on forests in northwestern North America. pp. 244-57, In: R.L. Peters and T.E. Lovejoy (Eds.), Global Warming and Biological Diversity, Yale University Press, New Haven, CT.

Schowalter, T.D. 1991. Roles of insects and diseases in sustaining forests. pp. 262-67 In: Society of American Foresters National Convention, 4-8 August 1991, San Francisco, CA. Society of American Foresters, Bethesda, MD.

Schowalter, T.D., B.A. Caldwell, S.E. Carpenter, R.P. Griffiths, M.E. Harmon, E.R. Ingham, R.G. Kelsey, J.D. Lattin and A.R. Moldenke. 1990. Decomposition of fallen trees: effects of initial conditions and heterotroph colonization rate. pp. 371-81, In: K.P. Singh, (Ed.) Ecological Management of Tropical Ecosystems, Wiley Eastern, Ltd., New Delhi.

Schowalter, T.D. 1990. Differences and consequences for insects. pp. 91-106, In: A.F. Pearson and D.A. Challenger (Eds.), Forests-Wild and Managed: Differences and Consequences. Students for Forestry Awareness, University of British Columbia, Vancouver, B.C.

Franklin, J.F., D.A. Perry, T.D. Schowalter, M.E. Harmon, A.McKee and T. Spies. 1989. Importance of ecological diversity in maintaining long-term site productivity. pp. 82-97, In: D.A. Perry, B. Thomas, R. Meurisse, R. Miller, J. Boyle, P. Sollins and J. Means (Eds.) Maintaining Long-term Productivity of Northwest Forests. Timber Press, Portland, OR.

Schowalter, T.D. and J.E. Means. 1989. Pests link site productivity to the landscape. pp. 248-50, In: D.A. Perry, B. Thomas, R. Meurisse, R. Miller, J. Boyle, P. Sollins, and J. Means, (Eds.) Maintaining Long-term Productivity of Pacific Northwest Forests. Timber Press, Portland, OR.

Schowalter, T.D. and J.M. Sexton. 1989. Inventory monitoring as a means of assessing insect impact on Douglas-fir seed production in western Oregon. pp. 151-60, In: Proc. 3rd Cone and Seed Insects Working Party Conference S2.07-01, Canadian Forestry Service, Pacific Forestry Centre, Victoria, B. C., Canada.

Carpenter, S.E., M.E. Harmon, E.R. Ingham, R.G. Kelsey, J.D. Lattin and T.D. Schowalter. 1988. Early patterns of heterotroph activity in conifer logs. In: R. Watling, A. Lyon and L. Boddy (Eds.) Fungi and Ecological Disturbance. Proc. Royal Soc. Edinburgh 94B:33-43.

Hansen, E.M., D.J. Goheen, P.F. Hessburg, J.J. Witcosky and T.D. Schowalter. 1988. Biology and management of black-stain root disease in Douglas-fir. pp. 63-80, In: T.C. Harrington and F.W. Cobb, Jr. (Eds.) Leptographium Root Diseases on Conifers. American Phytopathological Society Press, St. Paul, MN.

Crossley, D.A., Jr., C.S. Gist, W.W. Hargrove, Jr., L.S. Risley, T.D. Schowalter and T.R. Seastedt. 1987. Foliage consumption and nutrient dynamics in canopy insects. pp. 193-205, In: W.T. Swank and D.A. Crossley, Jr. (Eds.) Forest Hydrology and Ecology and Coweeta. Springer-Verlag, New York.

Schowalter, T.D. and D.A. Crossley, Jr. 1987. Canopy arthropods and their response to forest disturbance. pp. 207-18, In: W.T. Swank and D.A. Crossley, Jr. (Eds.) Forest Hydrology and Ecology at Coweeta. Springer-Verlag, New York.

Hansen, E.M., D.J. Goheen, P.F. Hessburg, J.J. Witcosky and T.D. Schowalter. 1986. Biology and management of black-stain root disease in Douglas-fir. pp. 13-19, In: O.T. Helgerson (Ed.) Forest Pest Management in Southwest Oregon. Forest Research Lab., Oregon State University, Corvallis OR.

Schowalter, T.D., M.I. Haverty, S.A. Dombrosky and J. Sexton. 1986. Response of Douglas-fir cone gall midge and Douglas-fir seed chalcid to host plant genotype. pp. 217-23, In: A. Roques, (Ed.), Proc. 2nd Cone and Seed Insects Working Party Conference S 2.07-01, Station de Zoologie Forestiére, INRA, CRF, Olivet, France.

Insect Ecology

    Insect population responses to environmental changes, such as natural disturbances (hurricances, drought and fire) and management practices (e.g., forest thinning and other harvest practices)

    Insect effects on primary production, decomposition of foliage and coarse woody debris, carbon and nutrient fluxes, and disturbance dynamics


Forest Entomology

    Cone and seed insects

    Conifer nursery insects

    Root-feeding beetles and vectored root pathogens

    Conifer aphids

    Defoliators

    Bark beetles and wood borers

B.A.     1974       Biology and Anthropology     Wichita State University

M.S.     1976       Biology                                        New Mexico State University

Ph.D.    1979       Entomology                                The University of Georgia