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I am interested in evolutionary and biogeographic patterns, and view systematic specialization as a means of examining such patterns on a manageable and personally satisfying scale. Most of my systematic research concentrates on the diversity and phylogenetic relationships of the staphylinid beetle subfamily Pselaphinae. Pselaphines possess an amazing array of species-level morphologies, including bizarre secondary sexual characters and sensory structures. They also exhibit highly derived evolutionary innovations associated with small size and a predatory lifestyle in interstitial spaces. As small organisms with extremely limited dispersal capabilities, many are indicators of relict faunal associations that reflect ancient biogeographic relationships. My other systematic interests extend in diverse directions, including some of the cucujoid families, especially Nitidulidae and Endomychidae. I particularly like the diversity of larval forms and developmental life histories within those groups.

In the field of conservation biology I am working towards a better understanding of the diversity and relationships of forest litter arthropod communities of natural ecosystems in the circum-Caribbean region. Current long-term projects include diversity studies in beech-magnolia forests on the Gulf Coastal Plain, and tropical forests in Central America and the Caribbean. Beech-magnolia forests in Louisiana occupy only a fraction of the area it covered prior to European settlement. This habitat is significant across the Gulf Coast because such forests harbor relict species of taxa that were displaced south during climatic changes that accompanied the advance of glaciers across much of eastern North America as recently as 15,000 years ago. The distributions of these species and their phylogenetic relationships provide important information for reconstructing historical events that had major impacts on the biota of the southern United States. Counterparts to these northern outliers include many species having affinities with neotropical taxa in Mesoamerica, particularly the Caribbean faunas of peninsular Florida, the West Indies, eastern Mexico, and the northern Central American countries of Belize, and Honduras. When sea levels dropped during Pleistocene glaciation, the land connections between these areas were much closer due to an extensive area of exposed continental shelf in the Gulf. Thus, it is important to consider these neighboring regions to develop a full understanding of the faunal associations of the Gulf Coast of the United States.

The current focus of broad scale research on beetle biodiversity is Great Smoky Mountains National Park along the Tennessee and North Carolina border. This is the most biologically diverse area in eastern United States thanks to its equable climate, its geographic location south of the limits of Pleistocene glaciation, and its mountainous terrain. The Park is the target of one of the most ambitious and long-standing All Taxa Biodiversity inventories in the world, initiated during the late nineties and is still going strong (see the Discover Life in America website). I'm in charge of coordinating Coleoptera studies for the project and pursue various specialized taxonomic projects within the context of the overall goals of the Coleoptera component of the ATBI. Our lab has been involved in the project 2001 and with acquisition of a major NSF Biotic Surveys and Inventory grant we will be able to continue to work for at least the next four years.

In addition to pursuing my own research program, I serve the agricultural community and the general public through identification and diagnosis of arthropods and related problems in Louisiana. As Director of the Lousiana State Arthropod Museum, I am responsible for fulfilling the objectives and managing the growth of the largest arthropod collection and research voucher repository in Louisiana. My teaching responsibilities include Insect Taxonomy and General Entomology (beginning 2006) courses.

Systematics of Coleoptera, especially the family Staphylinidae, subfamily Pselaphinae and the family Nitidulidae, biogeography, larval morphology, forest litter diversity and conservation biology.

Current Research Projects

Participant, Phylogenetic Relationships of Coleoptera Tree of Life project (in collaboration with NSF Co-PI’s B. Farrell, J. McHugh and M. Whiting).

Systematics and morphology of nitidulid beetle larvae (in collaboration with R. A. B. Leschen and A. R. Cline).

Revision of the neotropical pselaphine genera Barroeuplectoides and Bunoderus.

Beetle biodiversity in Great Smoky Mountains National Park (Leader, Taxonomic Working Group)

Systematics of the histerid beetle subfamily Hetaeriinae (in collaboration with A. K. Tishechkin).

Ecology of saproxylic Coleoptera (in collaboration with S. A. Gil and M. Ferro).

Pselaphine staphylinidae of New Zealand (in collaboration with D. S. Chandler and R. A. B. Leschen).

Specialized Taxonomic Activities

Identified/sorted Coleoptera for the following institutions (1995-2004): U.S. National Museum, Smithsonian Institution; American Museum of Natural History; Cornell University Insect Collection; Field Museum of Natural History; Great Smoky Mountains National Park Insect Collection; Snow Entomological Museum, University of Kansas; Essig Entomological Museum, University of California, Berkeley; Florida A & M University; Florida State Collection of Arthropods, University of Florida; Texas A & M University; Mississippi Entomological Museum, Mississippi State University; Montana State University; New Zealand National Collection of Athropods; Ohio State Museum of Biological Diversity, Ohio State University; Queensland Museum, Brisbane, Australia; Taiwan National University; Universidad De Catolica, Ecuador; University of Miami, Florida; University of New Mexico; U.S. Dept. of Agriculture.

Memberships in Professional Societies

American Entomological Society

Natural Science Collections Alliance (Institutional Membership)

Coleopterists Society

Entomological Collections Network

Entomological Society of America

European Association of Coleopterists

International Field Experience

Australia

1993, four weeks. Visiting specialist, Queensland Museum. Collecting trips to a variety of forest habitats in southern Queensland.

2005, two weeks. Collecting expedition to Northern Territory.

2007, two weeks. Collecting expedition to Western Australia.

Belize

1996, three weeks. Coleoptera survey of Rio Bravo Conservation Area.

1997, three weeks. Coleoptera survey of Rio Bravo Conservation Area and tropical insect biodiversity project development in collaboration with Programme for Belize and Belize National Plant Protection Service.

Chile

1994, seven weeks. North-south transect, collecting expedition in a variety of desert and forest habitats.

Costa Rica

1998, two weeks. Field research on Coleoptera associated with army ants at LaSelva Biological Station.

Ecuador

1991-1992, four weeks. Collecting expeditions to montane forests.

1990, three weeks. Collecting expeditions to montane forests.

1999, five weeks. Collecting expedition to Amazonian forest.

2008. five weeks. Collecting expedition to Amazonian forest.

Ghana

2001, two weeks. Collecting trip to Bobiri Forest Preserve, project development with Forestry Research Institute of Ghana.

India

2004, four weeks. Collecting trip to Assam and Megalaya.

New Zealand

1998, four weeks. Visiting specialist, Landcare Research. Collecting trip to South Island. Research in New Zealand National Collection of Arthropods.

2000, four weeks. Visiting specialist, Landcare Research. Collecting trip to North Island. Research in New Zealand National Collection of Arthropods.

2003-04, four weeks. Visiting specialist, Landcare Research. Collecting trip to North Island. Research in New Zealand National Collection of Arthropods.

2007, five weeks. Visiting specialist, Landcare Research. Collecting trip to South Island. Research in New Zealand National Collection of Arthropods.

Taiwan

2004, 2005, one week each; 2006, three weeks. Visiting specialist, Forestry Research Institute and Taiwan National University. Reconnaissance survey of field sites and meetings with Taiwanese collaborators.

Study at Major Museums

1979, one month; 2005, one week. Smithsonian Institution (USNM), Washington, DC.

1982, 1995, one week each. Field Museum of Natural History Chicago, IL.

1985, two days, American Museum of Natural History, New York, NY.

2000, one week. Alexander König Museum, Bonn, Germany

2000, three days. Canadian Museum of Nature, Ottawa, Canada.

2002, one week; 2008, two weeks. Natural History Museum, London, UK.

2002, one week. Muséum d'Histoire Naturelle, Geneva, Switzerland.

2008, Muséum national d'Histoire Naturelle, Paris, France.

Louisiana State University, Baton Rouge

1995-2000, Assistant Professor; 2000-2005, Associate Professor; 2005-2007, Professor, 2007-present, John Benjamin Holton Alumni Association Departmental Professorship in Agriculture, Department of Entomology. Research in insect systematics, Director, Louisiana State Arthropod Museum, teach systematics and morphology courses and advise graduate students.

University of Arkansas, Fayetteville

1989-1995: Research Associate, Department of Entomology. Conduct research in biodiversity and systematics, provide identifications of insects and diagnoses of related problems, and curate University of Arkansas Arthropod Museum.

1982-1989: Research Assistant (degree track), Department of Entomology. Manage entomology collection and provide insect identifications.

1977-1981: Graduate Assistant, Department of Entomology. Graduate student in Master's Program.

Grants and Contracts Awarded

2008

Sokolova, J., I. M. Sokolov, and C. E. Carlton. First survey of Microsporidia insect parasites in Great Smoky Mountains National Park. Discover Life in America. $4987.

2005

Carlton, C. E. Identification of beetles associated with course woody debris. USDA, Forest Service. $10,000.

Carlton, C. E. Long-horned and Leaf Beetles from the ATBI Structured Sampling and Publication of New Taxa for the Great Smoky Mountains National Park. Discover Life in America, Inc. $2500.

Carlton, C. E., and V. M. Bayless. Coleoptera of Great Smoky Mountains National Park. National Science Foundation. $651,054.

2004

Carlton, C. E., and V. M. Bayless. Acquisition of a High Resolution Digital Imaging System for the Department of Entomology. Louisiana Board of Regents Enhancement Program. $69,764.

2003

Carlton, C. E. Coleoptera Taxonomic Working Group (TWIG) Year 3: Identifying the catch. Discover Life in America. $4600.

2002

Carlton, C. E., and A. Cline. NSF Doctoral Dissertation Improvement Grant. The systematics and biogeography of Pocadius and related genera (Coleoptera: Nitidulidae). National Science Foundation. $3350.

Carlton, C. E. Continued support for the Coleoptera Taxonomic Working Group (TWIG) at the Louisiana State Arthropod Museum (LSAM). Discover Life in America. $6358.

Carlton, C. E., and L. Hooper-Bui. Screening Glassy-winged Sharpshooters for Egg Parasitoids in the Baton Rouge, LA Area. University of California, Riverside. $2000.

2001

Carlton, C. E. Coleoptera Taxonomic Working Group (TWIG) at the Louisiana State Arthropod Museum (LSAM). Discover Life in America. $4980.

2000

Carlton, C. E. Research experiences for undergraduates supplement. National Science Foundation. $5000.

1999

Weir, A., C. E. Carlton, and R. A. B. Leschen. South temperate beetles hangers: A survey of New Zealand Laboulbeniales fungi and their beetle hosts. National Science Foundation. $148,896.

Carlton, C. E., and A. K. Tishechkin. Systematics of the inquilinous hister beetle complex Mesynodites (Histeridae: Hetaeriinae). National Science Foundation. $81,300.

1997

Carlton, C. E., and V. L. Moseley. Expansion and enhancement of the Louisiana State Arthropod Museum. Louisiana Education Quality Support Fund. $56,000.

Carlton, C. E., and D. P. Prowell. Comparative insect diversity in natural ecosystems on the Gulf Coastal Plain. Louisiana Education Quality Support Fund. $89,000.

1996

Carlton, C. E. Glassy-winged sharpshooter, Homalodisca coagulata, egg parasitoid screening. University of California. $1000.

1995

Whitfield, J. B., and C. E. Carlton. Improvement of the University of Arkansas Arthropod Museum facilities. National Science Foundation. $51,920.

Carlton, C. E. Habitat transect survey of the American burying beetle in the Ouachita National Forest. United States Forest Service. $6000.

1994

Carlton, C. E. Annual monitoring survey of the American burying beetle on the Fort Smith, AR, Landfill (Coleoptera: Silphidae). City of Fort Smith. $27,500.

Carlton, C. E. Habitat transect survey of the American burying beetle in the Ouachita National Forest. United States Forest Service. $4500.

Carlton, C. E. Reconnaissance survey of terrestrial insects in the Cecil Creek area of the Buffalo National River. U. S. Department of Interior, National Parks Service. $3900.

Carlton, C. E. Biodiversity analysis of selected taxa from the Ouachita National Forest. United States Forest Service. $15,000.

1993

Carlton, C. E. Survey of the endangered American burying beetle, Nicrophorus americanus, in Arkansas (Coleoptera:Silphidae). Arkansas Game and Fish Commission. $25,000.

1992

Mangini, A, and Carlton, C. E. Preliminary survey of regeneration weevils and defoliators in the Ouachita National Forest. United States Forest Service. $14,992.

Carlton, C. E., and L. C. Thompson. Arthropod biodiversity study in the Ouachita National Forest. United States Forest Service. $44,000.

1991

Carlton, C. E. Survey of the American burying beetle, Nicrophorus americanus Oliver, in the Ouachita National Forest (Coleoptera: Silphidae). United States Forest Service. $1,000.

Carlton, C. E. Survey of substrate arthropods in deciduous climax forests in the Ouachita National Forest. United States Forest Service. $3,000.

1988

Carlton, C. E., and T. J. Kring. Biology and rearing requirements of Melitara prodenialis Walker (Lepidoptera: Phycitidae). Arkansas Science and Technology Authority. $13,595.

1987

Book Chapters

2009

Jelinek, J., C. E. Carlton, A. R. Cline, and R. A. B. Leschen. Nitidulidae Latrielle. In: Handbook of Zoology, Volume IV, Arthropoda, Insecta, part 38 (R. Beutel and R. A. B. Leschen, eds.). In press.

2003

Carlton, C. E. Revision of Reichenbachia of eastern North America (Coleoptera: Staphylinidae: Pselaphinae) [pp. 483-563]. In: R. Leschen and G. Cuccudoro (editors). Systematics of Coleoptera: Papers Celebrating the Retirement of Ivan Löbl, Associated Pub., Gainesville, FL. 955 pp.

Refereed Journal Articles

2009

Akbar, W., C. E. Carlton, and T. E. Reagan. Diomus terminatus (Say) (Coleoptera: Coccinellidae) life cycle, larval morphology, and potential as a biological control agent of the sugarcane aphid, Melanaphis sacchari Zehntner. Ann. Entomol. Amer.: 102: 96-103.

Carlton, C. E., and L. E. Watrous. First description of larvae in the tribe Mayetini: the larva of Mayetia pearsei (Coleoptera: Staphylinidae: Pselaphinae). Coleop. Bull.: in press.

Carlton, C. E., and R. A. B. Leschen. A new species and new North American record of Pseudotriphyllus (Coleoptera: Mycetophagidae). Coleop. Bull.: in press.

Ferro, M. L., Gimmel, M. L., and C. E. Carlton. The beetle community of small oak twigs in Louisiana with a review of literature featuring Coleoptera from fine woody debris. Coleop. Bull.: in press.

Robison, H. W., C. T. McAllister, C. E. Carlton, and G. Tucker. The Arkansas endemic biota: an update with additions and exclusion. Proc. Ark. Acad. Sci.: in press.

Watson, E. J., J. P. Geagan, and C. E. Carlton. Biodiversity of necrophilous insect communities associated with wildlife carcasses using pitfall trap data. Florida Entomol.: submitted.

2008

Carlton, C. E. Eight new species of Arianops Brendel from southeastern United States with an updated key and notes on additional species (Coleoptera: Staphylinidae: Pselaphinae). Coleop. Bull. 62: 297-32.

Carlton, C. E., and R. A. B. Leschen. Description of the larva of Pselaphophus atriventris (Westwood) (Staphylinidae: Pselaphinae: Pselaphini) with notes on its life history and a list of described pselaphine immature stages. Ann. Entomol. Soc. Amer. 101: 13-19.

Gimmel, M. L., and C. E. Carlton. The identity of Languria femoralis Motschulsky (Erotylidae: Languriinae). Zootaxa 1821: 67-68.

Radke, M. G., C. E. Carlton, and G. B. Williamson. A Dung Beetle Assemblage in an Urban Park in Louisiana. Southeastern Naturalist 7: 101-110.

Sokolov, I. M., and C. E. Carlton. Two new species of blind, forest litter-inhabitiing ground beetles from the subtribe Anillina (Carabidae: Trechinae: Bembidiini) from eastern U.S.A. Zootaxa 1740: 37-44.

2007

Carlton, C. E., and V. M. Bayless. Documenting beetle diversity in Great Smoky Mountains National Park; beyond the halfway point! (Arthropoda: Insecta: Coleoptera). Southeastern Naturalist Special Publication 1: 183-192.

Carlton, C. E., and R. A. B. Leschen. Descriptions of Soronia (Coleoptera: Nitidulidae) larvae of New Zealand with comments on life history and taxonomy. New Zealand J. Zool. 30: 41-51.

Reay-Jones, F. P. F., L. T. Wilson, M. O. Way, C. E. Carlton and T. E. Reagan. Movement of the Mexican rice borer through the Texas rice belt. J. Econ. Entomol. 100: 54-60.

Sokolov, I. M., Y. Y. Sokolova, and C. E. Carlton. New species of Anillinus casey (Carabidae: Trechinae: Bembidiini) from Great Smoky Mountains National Park, U.S.A. and phylogeography of the A. langdoni species group. Zootaxa 1542: 1-20.

2005

Bayless, V. M., and C. E. Carlton. Sifting Litter in the Great Smoky Mountains. Southeastern Biol. 52: 435-436.

McAllister, C. T., and C. E. Carlton. Second record of the dipluran, Occasjapyx carltoni Allen, 1988 (Japygidae) from Arkansas. Proc. Ark. Acad. Sci. 59: 213.

Watson, E. J., and C. E. Carlton. Insect succession and decomposition of wildlife carcasses during fall and winter in Louisiana. J. Med. Entomol. 42: 193-203.

Watson, E.J. and C.E. Carlton. Succession of forensically important Silphidae species on large carcasses (Coleoptera). Southeastern Naturalist 4: 335-346.

2004

Carlton, C. E., M. Dean, and A. K.Tishechkin. Diversity of two beetle taxa at a Western Amazonian locality (Coleoptera: Histeridae; Staphylinidae, Pselaphinae). Coleop. Bull. 58: 163-170.

Carlton, C. E., and R. S. Anderson. Occurrence of the Introduced Weevil Myosides seriehispidus Roelofs in Great Smoky Mountains National Park (Coleoptera: Curculionidae). Coleop. Bull. 58: 343.

Cline, A. R., and C. E. Carlton. Two new species of Epuraea (Orthopeplus) from Mexico (Coleoptera: Nitidulidae). Coleop. Bull. 58: 261-270.

Cline, A. R., and C. E. Carlton. Three new species of Lasiodactylus Perty from the Neotropics (Coleoptera: Nitidulidae: Nitidulinae). Coleop. Bull. 58: 355-368.

Leschen, R. A. B., and C. E. Carlton. A New Tribe, Genus and Species of Nitidulid Beetle (Coleoptera: Nitidulidae: Nitidulinae) from Bolivia. Coleop. Bull. 58: 443-451.

Sokolov, I. M., C. E. Carlton, and J. F. Cornell. Review of Anillinus with descriptions of 13 new species and a key to soil and litter species (Coleoptera: Trechinae: Bembidiini). Coleop. Bull. 58: 185-233.

2003

Cox, R. T., and C. E. Carlton. A comment on gene introgression versus en masse cycle switching in the
evolution of 13-year and 17-year life cycles in periodical cicadas. Evolution 57: 428-432.

Van Zandt, P. A., V. R. Townsend, Jr., C. E. Carlton, M. Blackwell, and S. Mopper. Loberus impressus (Coleoptera, Erotylidae) fungal associations and presence in the seed capsules of Iris hexagona. Coleop. Bull. 57: 281-288.

Watson, E. J., and C. E. Carlton. Spring Succession of Necrophilus Insects of Wildlife Carcasses in Louisiana. J. Med. Entomol. 40: 338-347.

2002

Bossart, J. L., and C. E. Carlton. Insect Conservation in America: status and perspectives. Amer. Entomol. 48(2): 82-92.

2001

Carlton, C. E., and R. A. B. Leschen. Species in sympatry: Pselaphotheseus of Campbell Island (Coleoptera: Staphylinidae: Pselaphinae). New Zealand J. Zool. 28: 387-393.

2000

Carlton, C. E., V. R. Townsend, Jr., P. A. Van Zandt, and S. Mopper. Description of the larva of Loberus impressus LeConte with notes on its natural history (Coleoptera: Languriidae: Xenoscelinae). Ann. Entomol. Soc. Amer. 93: 356-361.

Leschen, R. A. B., and C. E. Carlton. A new species of Micropsephodes from southern United States (Endomychidae: Anamorphinae). Coleop. Bull. 54: 232-238.

1999

Landau, D., D. P. Prowell, and C. E. Carlton. Intensive versus long-term sampling to assess lepidopteran diversity in a southern mixed mesophytic forest. Ann. Entomol. Soc. Amer. 92: 435-441.

1998

Carlton, C. E., and H. W. Robison. Diversity of litter-dwelling beetles in deciduous forests of the Ouachita highlands of Arkansas (Insecta: Coleoptera). Biodiv. and Cons. 7: 1589-1605.

Carlton, C. E., and F. Rothwein. The endangered American burying beetle (Coleoptera: Silphidae: Nicrophorus americanus at the edge of its range in Arkansas. Coleop. Bull. 52: 179-185.

Cox, R. T., and C. E. Carlton. A commentary on prime numbers and periodical cicadas. Amer. Nat. 152: 162-164.

Meek, C. L., C. Puskarich-May, and C. E. Carlton. New state record for the hairy maggot blow fly Chrysomya rufifacies (Macquart). Southwestern Entomol. 23: 373-375.

Muegge, M. A., and C. E. Carlton. New records of Japygoidea from Louisiana with notes on behavior (Hexapoda: Diplura). Entomol. News 109: 177-182.

Triapitsyn, S. V., R. F. Mizell, III, J. L. Bossart, and C. E. Carlton. Egg Parasitoids of Homalodisca coagulata (Homoptera: Cicadellidae). Florida Entomol. 81: 241-243.

1997

Robison, H. W., G. W. Byers, and C. E. Carlton. An annotated checklist of the Mecoptera (Scorpionflies) of Arkansas. Entomol. News. 108: 313-317.

1996

Carlton, C. E., and Lori Spencer Nobles. Distribution of Speyeria diana in the highlands of Arkansas, Missouri and Oklahoma with comments on conservation. Entomol. News. 107: 213-219.

Carlton, C. E., and H. W. Robison. Notes on Mayetia pearsei Schuster, Marsh and Park with a revised key to eastern Mayetia species (Staphylinidae: Pselaphinae). Coleop. Bull. 50: 244-250.

Carlton, C. E., and R. A. B. Leschen. New species, distributional notes and systematics of Eutrichites (Coleoptera: Staphylinidae: Pselaphinae). Trans. Amer. Entomol. Soc. 122: 161-173.

Carlton, C. E., R. A. B. Leschen, and P. L. Kovarik. Adult blow fly predation in a Chilean Euspilotus (Coleoptera: Histeridae). Coleop. Bull. 50: 154.

Dorris, P. R., H. W. Robison, and C. E. Carlton. Spiders (Arthropoda: Aranea) from deciduous forest litter of the Ouachita Highlands. Proc. Ark. Acad. Sci. 49: 45-48.

Leschen, R. A. B, and C. E. Carlton. Slime production in mycophagous Nitidulidae (Coleoptera) including a new species of Eusphaerius. J. of Nat. Hist. 30: 1861-1873.

C. S. Martin, C. E. Carlton, and C. L. Meek. New distribution record for the hairy maggot blow fly Chrysomya rufifacies. Southwestern Entomol. 21: 477-478.

1995

Carlton, C. E. A new species of Pseudactium from Arkansas (Coleoptera: Staphylinidae: Pselaphinae). Coleop. Bull. 49: 170-173.

Carlton, C. E., and J. L. Lancaster. Arkansas horse and deerflies (Insecta: Diptera: Tabanidae). Ark. Agri. Exp. Sta. Bull. Ser. 27 pp.

1994

Carlton, C. E., and D. S. Chandler. Revision of the nearctic genus Pseudactium (Coleoptera: Pselaphidae: Euplectinae). Coleop. Bull. 48: 171-190.

Carlton, C. E., and T. J. Kring. Melitara prodenialis Walker on prickly pear in Arkansas. Southwestern Entomol. 19: 23-31.

Phillips, E. C., R. V. Kilambi, and C. E. Carlton. Life history and secondary production of Ephoron album (Say) (Ephemeroptera: Polymitarcidae) in the Illinois River, Arkansas. J. Kansas Entomol. Soc.: 67: 242-247.

1993

Leschen, R. A. B., and C. E. Carlton. Three new species and a new record of neotropical Pocadius Erichson (Coleoptera: Nitidulidae). Trop. Biol. 7: 209-216.

Leschen, R. A. B., and C. E. Carlton. Debris cloaking in Endomychidae: A new species from Peru. J. Zool. (Linnean Soc.) 109: 35-51.

Springer, T. L., and C. E. Carlton. Dark-winged fungus gnat (Diptera: Sciaridae) ovipositional preference for Trifolium species. J. Econ. Entomol. 86: 1420-1423.

Hopkins, J. D., C. D. Steelman, and C. E. Carlton. Internal reproductive system of the adult male lesser mealworm Alphitobius diaperinus (Coleoptera: Tenebrionidae). J. Kansas Entomol. Soc. 66: 446-450.

1992

Hopkins, J. D., C. D. Steelman, and C. E. Carlton. Anatomy of the adult female lesser mealworm, Alphitobius diaperinus (Coleoptera: Tenebrionidae) reproductive system. J. Kansas Entomol. Soc. 65: 299-307.

1991

Cox, R. T., and C. E. Carlton. Evidence for genetic dominance in 13-year periodical cicadas (Homoptera: Cicadidae: Magicicada spp.). J. Amer. Midl. Nat. 125: 63-74.

1990

Carlton, C. E., and R. T. Cox. A new species of Arianops from central Arkansas and the biogeographic implications of the Interior Highland Arianops species (Coleoptera: Pselaphidae). Coleop. Bull. 44: 365-371.

Carlton, C. E. Biogeographic affinities of pselaphid beetle genera of the Eastern United States. Florida Entomol. 73: 570-579.

1989

Allen, R. T., and C. E. Carlton. New records of Schizopteridae and Ceratocombidae (Hemiptera: Dipsocoroidea). J. Kansas Entomol. Soc. 62: 125-126.

Carlton, C. E. Revision of the genus Eutrichites (Coleoptera: Pselaphidae). Coleop. Bull. 43: 105-119.

Carlton, C. E. Aedeagal isomerism in Actizona chuskae Chandler (Coleoptera: Pselaphidae) and a new locality record. Coleop. Bull. 43: 57-58.

Carlton, C. E., and R. T. Allen. A new species of Arianops (Coleoptera: Pselaphidae) and the rediscovery of Arianops sandersoni Barr. Coleop. Bull. 43: 59-67.

Carlton, C. E., and E. H. Schmitz. Anatomy of the extrinsic gut musculature of Gammarus minus (Crustacea: Amphipoda). J. of Morphol. 200: 87-92.

1988

Allen, R. T., and C. E. Carlton. Two new species of Scaphinotus (Coleoptera: Carabidae), with comments on other Arkansas species. J. New York Entomol. Soc. 96: 129-139.

Allen, R. T., C. E. Carlton, and S. A. Tedder. A new species of Acalypta (Hemiptera: Tingidae). J. Kansas Entomol. Soc. 61: 126-130.

Carlton, C. E. Dienerella filum (Aube), a potential pest of air conditioning systems (Coleoptera: Lathridiidae). Coleop. Bull. 42: 263-264.

Carlton, C. E., and D. G. Bassi. Oligotoma nigra (Embiidina: Oligotomidae), an introduced species new to Arkansas. J. Kansas Entomol. Soc. 61: 352-353.

Cox, R. T., and C. E. Carlton. Paleoclimatic influences in the evolution of periodical cicadas (Homoptera: Cicadidae: Magicicada spp.). J. Amer. Midl. Nat. 120: 183-193.

Leschen, R. A. B., and C. E. Carlton. Immature stages of Endomychus biguttatus Say (Coleoptera: Endomychidae), with observations on the alimentary canal. J. Kansas Entomol. Soc. 61: 321-327.

1986

Carlton, C. E., and R. T. Allen. Revision of the genus Euboarhexius (Coleoptera: Pselaphidae). Coleop. Bull. 40: 285-296.

1983

Carlton, C. E. Revision of the genus Conoplectus Brendel (Coleoptera: Pselaphidae). Coleop. Bull. 37: 55-80.

Non-refereed Articles and other Contributions

2008

Carlton, C. E. Musings on sabbaticals and beach scrabblers. The Entomological Society of New Zealand Newsletter. January 2008: 2-3.

Hummel, N., D. Pollet, G. Reagan, W. Akbar, J. Beuzelin, C. Carlton, J. Saichuk, and M. Way. Mexican Rice Borer (Eoreuma loftini Dyar). LSU AgCenter Publication 3098, 2 pp.

2007

Carlton, C. E. Book Review. Coleoptera, Beetles. Volume 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). Handbook of Zoology. Insecta. Part 38 (Beutel, R. D., and R. A. B. Leschen, eds.). Coleop. Bulletin 61: 492-493.

2005

Carlton, C. E., D. S. Chandler, R. A. B. Leschen, E. G. Riley, and P. E. Skelley. Obituary and Dedication. Karl Heinz Stephan 1931-2005. Coleop. Bull. 59: 277-283.

Carlton, C. E. Book Review. American Beetles Volume I. Archostemata, Myxophaga, Polyphaga: Staphyliniformia. American Beetles Volume II. Polyphaga: Scarabaeoidea through Curculionoidea. Coleop. Bull. 59:154-156.

2004

Nash, S. Desperately Seeking Charisma: Improving the Status of Invertebrates. Bioscience 54: 487-494 (published color photograph, p. 488).

2002

Carlton, C. E. Mining Beetle Diversity in the Smokies. Southeastern Biology 49: 27-28.

2001

Carlton, C. E. Coleopterists Society Monographs, Patricia Vaurie Memoir Series. Coleop. Bull. 55: 528-529.

1994

Carlton, C. E., J. Bollinger, and L. C. Thompson. Arthropod biodiversity-sampling protocol development. Proceedings of the Symposium on Ecosystem Management Research in the Ouachita Mountains: Pretreatment Conditions and Preliminary Findings. pp 144-153.

Mangini, A., C. Carlton, R. W. Perry, and J. Hanula. Seed, cone, regeneration, and defoliating insects in forest ecosystem management. Proceedings of the Symposium on Ecosystem Management Research in the Ouachita Mountains: Pretreatment Conditions and Preliminary Findings. pp 154-161.

1987

Carlton, C. E. Identification of Arkansas fire ant (Hymenoptera: Formicidae: Solenopsis spp.) workers. Ark. Agri. Exp. Stat. Rep. Ser. 301. 12 pp.

1986

Carlton, C. E. Fossil collecting in Arkansas. Part 2. The Arkansas Naturalist 4, no. 4, pp. 7-26.

1985

Carlton, C. E. Arkansas museums. The Arkansas Naturalist 3, no. 4, pp. 4-14.

Carlton, C. E. Fossil collecting in Arkansas. Part 1. The Arkansas Naturalist 3, no. 8, pp. 1-16.

1984

Carlton, C. E. Denizens of the duff. The Arkansas Naturalist 2, no. 1, pp. 6-15.

Carlton, C. E. Checklist of the longhorn beetles of Arkansas (Coleoptera: Cerambycidae). Ark. Acad. Sci. Ark. Biota Survey, Checklist No. 40. 5 pp.

Academic Publications Edited

2005

Aalbu, R. L., F. G. Andrews, and D. A. Pollock. Revision of Western Hemisphere Dacoderinae with a Phylogenetic Analysis of World Taxa (Coleoptera: Salpingidae). Coleopterists Society Monographs Patricia Vaurie Series No. 3. 42 pp.

Leschen, R. A. B., and I. Löbl. Phylogeny and Classification of Scaphisomatini (Staphylinidae: Scaphidiinae) with Notes on Mycophagy, Termitophily, and Functional Morphology. Coleopterists Society Monographs Patricia Vaurie Series No. 4. 62 pp.

2004

Clark, S. M., D. G. LeDoux, T. N. Seeno, E. G. Riley, A. J. Gilbert, and J. S. Sullivan. Host Plants of Leaf Beetle Species Occurring in the United States and Canada (Coleoptera: Megapodidae, Orsodacnidae, Chrysomelidae, excluding Bruchinae). The Coleopterists Society Special Publication No. 2. 476 pp.

2003

Paucar-Cabrera, A. Systematics and Phylogeny of the Genus Epectinaspis Blanchard (Coleoptera: Scarabaeidae: Rutelinae) and Description of a New Genus of Anomalini from Mexico. Coleopterists Society Monographs Patricia Vaurie Series No. 2. 60 pp.

Riley, E. G., S. Clark, and T. Seeno. Catalog of the Leaf Beetles of America North of Mexico. The Coleopterists Society Special Publication No. 1. 290 pp.+search and sort CD.

2002

Cook, J. Revision of the Neotropical Genus Cryptocanthon Balthasar (Coleoptera: Scarabaeidae: Scarabaeinae). Coleopterists Society Monographs Patricia Vaurie Series No. 1. 93 pp.

1999-2005

Coleopterists Bulletin 53(4), 96 pp; 54 (1-4), 552 pp; 55 (1-4), 528 pp.; 56 (1-4), 608 pp.; 57 (1-4), 510 pp.; 58 (1-4) 680 pp; 59 (1-3) 405 pp.

Electronic Publications

2007

Foil, L. D., C. E. Carlton, and W. L. Kramer. Chagas disease in Louisiana. LSU AgCenter online publication. http://www.lsuagcenter.com/en/environment/insects/Chagas+Disease+in+Louisiana.htm

2004

Carlton, C. E. Current Status of Nitidulid Beetle Larval Taxonomy. No 6 on CD Rom, Phylogeny of the Cucujoidea (F. W. Shockley, producer). University of Georgia, Dept. of Entomology.

2001-present

Carlton, C. E. Great Smoky Mountain National Park Coleoptera ATBI TWIG Website. Http://entomology.lsu.edu/lsam/smokybeetles.htm

1998-present

Carlton, C. E. The Louisiana State Arthropod Museum Website. Http:// entomology.lsu.edu /lsam/index.htm

1998-2007

Management of the Department of Entomology Website. Http://www.lsu.edu.departments.entomology.htm

2001-2005

Carlton, C. E. Coleopterists Bulletin Manuscript Preparation Guidelines. Http://www.coleopsoc.org/coleguid.shtml

Presentations at Professional Meetings, Symposia, Seminars, and Conferences

2007

Ferro, M. L., M. L. Gimmel, and C. E. Carlton. Fine beetles: diversity of twig bundles and efficacy of using them as a collecting technique. Poster presentation. Entomological Society of America Annual Meeting, San Diego, CA.

2006

Bayless, V. M., and C. E. Carlton. The Louisiana State Arthropod Museum. Invited presentation. Entomological Collections Network Annual Meeting, Indianapolis, IN.

Carlton, C. E. Beetle discoveries and diversity in Great Smoky Mountians National Park. Symposium presentation. Entomological Society of America Annual Meeting, Indianapolis, IN.

Carlton, C. E., and V. M. Bayless. Documenting beetle diversity in Great Smoky Mountains National Park; beyond the halfway point. Invited presentation. Association of Southeastern Biologists Annual Conference, Gatlinburg, TN.

Ferro, M. L., and C. E. Carlton. Saproxylic beetles in the rough. Poster presentation. Entomological Society of America Annual Meeting, Indianapolis, IN.

Gimmel, M. L., and C. E. Carlton. Confronting the taxonomic impediment from behind: making old coleopteran literature freely available. Poster presentation. Entomological Society of America Annual Meeting, Indianapolis, IN.

Ratcliffe, B. C., Carlton, C. E. (co-organizers). Notes from the Field: Biodiversity of Coleoptera. A Symposium Honoring the Memory and Contributions of Steve Ashe. Parts 1 and 2. Section A Symposia. Entomological Society of America Annual Meeting, Indianapolis, IN.

2005

Carlton, C. E. Pselaphine Staphylinids of Great Smoky Mountains National Park (Coleoptera: Staphylinidae: Pselaphinae). Display presentation. Discover Life in American Annual Conference, Gatlinburg, TN and Entomological Society of America Annual Meeting, Ft. Lauderdale, FL.

Carlton, C. E., and V. M. Bayless. Coleoptera of the Great Smoky Mountains National Park. Display presentation. Discover Life in American Annual Conference, Gatlinburg, TN and Entomological Society of America Annual Meeting, Ft. Lauderdale, FL.

Cline, A. R., and C. E. Carlton. The Sap Beetles of the Great Smoky Mountains National Park (Coleoptera: Nitidulidae). Discover Life in American Annual Conference, Gatlinburg, TN and Entomological Society of America Annual Meeting, Ft. Lauderdale, FL.

Ferro, M., C. E. Carlton, and K. Klepzig. Effect of Loblolly Pine Coarse Woody Debris (CWD) on Coleoptera Species Richness. Display presentation. Entomological Society of America Annual Meeting, Ft. Lauderdale, FL.

Gimmel, M., C. E. Carlton, and A. J. Mayor. Scirtidae of Great Smoky Mountains National Park (Coleoptera: Scirtidae). Display presentation. Discover Life in American Annual Conference, Gatlinburg, TN and Entomological Society of America Annual Meeting, Ft. Lauderdale, FL.

McAllister, C. T., and C. E. Carlton. Second Record of the Dipluran, Occasjapyx carltoni Allen, 1988 (Insecta: Japygidae), from Arkansas. Arkansas Academy of Sciences Annual Meeting, Conway, AR.

Reay-Jones, P. F., M. O. Way, L. T. Wilson, T. E. Reagan, and C. E. Carlton. Monitoring the Movement of the Mexican Rice Borer (Lepidoptera: Crambidae) with Pheromone Baited Traps. Oral presentation. Entomological Society of America Annual Meeting, Ft. Lauderdale, FL.

Tindall, K., F. Huang, C. E. Carlton, and V. M. Bayless. Moodna bisinuella Hampson: A New State Record for Louisiana and a Potential Threat to Corn. Display presentation. Entomological Society of America Annual Meeting, Ft. Lauderdale, FL.

2004

Cline, A. R., and C. E. Carlton. Evolution of the Egg Laying Apparatus in the Pocadiini (Coleoptera: Nitidulidae). Display presentation. Entomological Society of America Annual Meeting, Salt Lake City, UT.

Carlton, C. E. The Great Smoky Mountain All Taxa Biodiversity Inventory: Combining Science, Education, and Fun. Weekly Seminar Series, LSU Museum of Natural Science, Baton Rouge, LA, April 2004.

2003

Carlton, C. E. Coleoptera Twig Status Report: Year 3. Annual Conference, Discover Life in America, Gatlinburg, TN.

Carlton, C. E. Current Status of Nitidulid Beetle Larval Taxonomy. Invited presentation. Section A Symposium, Annual Meeting, Entomological Society of America, Cincinnati, OH.

Cline, A. R., C. E. Carlton, and J. L. McHugh (coorganizers). Phylogeny of the Cucujoidea I and II. Section A Symposia, Annual Meeting, Entomological Society of America, Cincinnati, OH.

2002

Carlton, C. E. Coleoptera Twig Status Report: Year 2. Annual Conference, Discover Life in America, Gatlinburg, TN.

Carlton, C. E. Diversity of forest litter inhabiting Coleoptera and other wonders. Departmental Seminar, Department of Entomology, Louisiana State University, Baton Rouge, LA.

Reagan, T. E., M. O. Way, C. E. Carlton, and F. R. Posey. Update on Mexican Rice Borer Research for the Rice and Sugar Cane Industries of Texas and Louisiana. Display presentation. Entomological Society of America Annual Meeting, Fort Lauderdale, FL.

2001

Carlton, C. E. Status of the Coleoptera TWIG of the Great Smoky Mountain National Park ATBI. Annual Conference, Discover Life in America, Gatlinburg, TN.

Carlton, C. E., and J. L. Bossart. The prejudice and promise of insect conservation research in the United States. Southeastern Branch, Entomological Society of America, Augusta, GA.

Carlton, C. E. Introduction to pselaphidology and the world of litter arthropods. Departmental Seminar, Department of Biological Sciences, Louisiana State University, Baton Rouge, LA.

2000

Carlton, C. E. Phylogenetic relationships among major lineages of pselaphine staphylinids (Coleoptera: Staphylinidae). Invited presentation. Section A Symposium, Annual Meeting, Entomological Society of America, Montreal, Canada.

Carlton, C. E., M. K. Dean, and A. K. Tishechkin. Diversity of two beetle taxa at a western Amazonian locality. Poster presentation. XXI International Congress of Entomology, Iquassu Falls, Brazil.

Reagan, T. E., C. E. Carlton, and F. R. Posey. The Mexican rice borer threat to sugarcane and rice in Texas and western Louisiana. Poster presentation. Entomological Society of America, Montreal, Canada and Southwestern Branch Meeting, ESA, Austin, TX.

1999

Carlton, C. E. Diversity and biogeographic significance of endemic and disjunct beetles of the Ouachita Mountains. Invited presentation. Symposium on Ecosystem Management Research in the Ouachita and Ozark Mountains. Hot Springs, AR.

Carlton, C. E. The public service role of Land Grant University-based entomology museums. Student sponsored symposium presentation. Annual Meeting, Southeastern Branch, Entomological Society of America. Destin, FL.

Carlton, C. E. Diversity and biogeography of forest litter Coleoptera in the Southeastern United States. Departmental seminar, Department of Biology, Southwestern Louisiana State University, Layfayette, LA.

1998

Carlton, C. E. What arcane taxa tell us about the way the world ought to be. Charles Lincoln Distinguished Lecture Series. Department of Entomology, University of Arkansas, Fayetteville.

Tishechkin, A. K., and C. E. Carlton. Mesynodites s. l.: A barrier to understanding phylogenetic relationships within the Hetaeriinae. Display presentation, Annual Meeting, Entomological Society of America, Las Vegas, NV.

Wheeler, A. F., Jr., C. E. Carlton, and D. Landau (Co-organizers). Arthropods of unique communities and habitats in the southeastern United States. Symposium, Southeastern Branch Meeting, Entomological Society of America, Chattanooga, TN.

Carlton, C. E. Insects and the conservation of mixed mesophytic forest habitat on the Gulf Coastal Plain. Symposium presentation, Southeastern Branch Meeting, Entomological Society of America, Chattanooga, TN.

1997

Carlton, C. E. Patterns of endemism on the Gulf Coastal Plain in pselaphine staphylinids (Coleoptera: Staphylinidae: Pselaphinae). Oral presentation. Southeastern Branch Meeting, Entomological Society of America, Asheville, NC.

1995

Carlton, C. E. The genus Reichenbachia in eastern North America (Coleoptera: Pselaphidae). Oral presentation. Annual Meeting, Entomological Society of America, Las Vegas, NV.

Carlton, C. E. Silphid research in Arkansas. Conference on the American Burying Beetle. Invited paper, Norman, OK.

Carlton, C. E. Chile, a wrath of grapes. Departmental seminar, University of Arkansas, Fayetteville, AR.

Carlton, C. E. Introduction to pselaphine Staphylinidae. Invited Seminar, Louisiana State University, Department of Entomology, Baton Rouge, LA.

1993

Carlton, C. E. Arthropod biodiversity-sampling protocol development. Invited paper, U. S. Forest Service Symposium on Ecosystem Management, Hot Springs, AR.

Carlton, C. E. Forest substrate biodiversity research in the Ouachita Mountains. Departmental seminar, University of Arkansas, Fayetteville, AR.

1992

Carlton, C. E. Coleoptera diversity in deciduous forest substrates in the Ouachita Highlands of Arkansas. Display presentation, Annual Meeting, Entomological Society of America, Baltimore, MD.

Carlton, C. E., and R. A. Leschen. Methodology and theory in insect systematics. Iinvited seminar, Biology Department, University of Azuay, Cuenca, Ecuador.

1990

Carlton, C. E., and R. A. Leschen. Debris cloaking in Bystus “decorator” (Coleoptera: Endomychidae) larvae and the morphology of the thoracic musculature. Oral presentation. Annual Meeting, Entomological Society of America, New Orleans, LA.

1989

Biogeographic affinities of pselaphid beetles of the eastern United States. Symposium presentation, Southeastern Branch Meeting, Entomological Society of America, Orlando, FL.

1988

Carlton, C. E. Life history of the cactus boring moth Melitara prodenialis in Arkansas (Lepidoptera: Pyralidae). Oral presentation. Annual Meeting, Entomological Society of America, Louisville, KY.